By
now we have a good understanding of the structural anatomy of the GSD
and its basic gaits. Now we must turn our attention to how the dog moves
at an anatomical level, to give us a good understanding of why some
kinds of structure are desirable and others are not. I think the most
logical approach is to follow the progression of energy, from the first
generation of power, along its transmission to its final dissipation
(Fig 1). The less fragmented this line of progression, the more concentrated
the power and the more efficient the dogs movement. Faults of
structure result in energy leakage, and the more leakage, the less is
available for the wanted task. Leakage also creates stress on areas
of anatomy in ways they are not designed to withstand. For the sake
of convenience, and because it is the gait at which the breed is commonly
judged, well focus initially on the trot, discussing aspects of
the other gaits where appropriate.

First
however, a very brief anatomy lesson. The leg consists of several components.
The bones are the underlying architecture that give support, strength
and basic structure. To them are attached the muscles, which for our
purposes fall into two groups, flexors and extensors. Flexors are those
muscles that, when they contract, close the joint and flex the limb,
as when you lift your leg or crook your arm (Fig 2). Extensors, when
they contract, do the opposite, opening the joint and straightening
the limb (Fig 3). Muscles are fleshy masses which at their ends become
tendons: tough, stringy fibers that act as ropes upon which the muscle
will pull to move a bone. The tip of the tendon is attached to a process,
a bony protuberance that gives the muscle extra leverage in moving the
bone.
For instance, in both dogs and humans the
calf muscle is a large muscle attached at the top of the back of the
lower leg bone, the fibia, and that stretches down to become the Achilles
tendon at the back of the ankle, attaching to the bony process that
is the heel bone (Fig 4). When the calf muscle contracts, it shortens
dramatically and powerfully, pulling the heel bone towards it. The foot
is like a seesaw and pivots at the ankle joint, the heel going up and
the toes going down. In humans this muscle is so powerful that with
just one leg it can lift and propel the entire weight of a large man.
Some muscles are situated on the inside of a joint and span the angle
between two bones to pull them together and bend the joint. Muscles
at the back of the thigh reach down and attach to the bone below the
knee, and when they contract, the knee (or stifle) bends (Fig 5). Other
muscles reach around the outside of a joint. The front thigh muscle
sends its tendon over the top of the knee to attach to the top of the
shin. When it contracts, it pulls the bones into alignment and straightens
the knee (Fig 6).



A
muscles power is related to its shape. When relaxed, it is soft
and has no strength. It generates power by contracting, or shortening
lengthwise, and becoming denser and hard to the touch. A long, slim,
flat muscle can contract to a huge degree, losing a third of its length
and giving the bone to which it is attached a great deal of leverage
and a wide range of movement. That is why gazehounds built for speed
have long, slim, flat muscling. Short thick muscles cannot shorten so
dramatically, but because they are so well endowed with muscle fibres
their contractions are very powerful and sustained. They cant
provide sweeping movement, but they do offer great strength. So malamutes,
mastiff and bully breeds, who require strength and endurance, have short,
thick muscles (Fig 7a).
Similarly,
the bones reflect the requirements of speed or strength. Long bones
give great leverage and long strides for speed and jumping ability.
However, they are usually lighter and can be more fragile. Antelope
and deer, even the cheetah, are ideal examples. Short, thick bones cant
give exceptional leverage, but they have weight, strength and stability,
and can withstand the stress generated by massive muscles. Badgers,
wolverines, bear and bulldogs are typical examples (Fig 7b).
The GSD must have a balance of speed, agility,
strength and endurance. He must be able to outrun and out jump a young
man, subdue an angry 200lb ram and patrol a moving flock all day, every
day. A tall, thin, light dog may not have the power to protect himself
or his handler in a serious fight. The heavy, low stationed, short legged
dog will not have the stride to gallop or jump well, and his weight
will sap his energy. This breed has to have it all, but without one
characteristic becoming so developed as to atrophy another. Certainly,
fitness and conditioning can go a long way to compensate a dog's shortcomings,
but having good structure in the first place, plus conditioning, will
give the dog the best possible advantage whatever its task.
Power
generation begins in the rear. It starts at that point in the stride
where the hind foot takes on the bodys full weight and can begin
to drive it forward (Fig 8). This happens when the paw is planted vertically
beneath the hip joint. In this position, the leg is at maximum contraction.
This is what we call the support position. During the phase where the
paw is ahead of the hip at the forward part of the stride it is beginning
to support the body weight, but is still contracting. There is some
contraction of the extensor muscles, sufficient to keep the body supported,
but full extension at this stage would push the body backwards. Obviously,
there is some upward propulsion as well. At the beginning of propulsion,
it is nearly straight up, and as the stride progresses, the line of
propulsion up the leg slants more and more forward. Some upward force
at the beginning of the stride is desirable. It gives the body that
bit of lift required for the period of suspension when the body travels
forward without contact with the ground, purely of its own momentum.
Momentum gives motion without effort, so a long period of suspension
is highly desirable. Too much lift however, can make the dog appear
to bounce up and down and can actually shorten the period of suspension,
wasting energy.
At
a walk, very little propulsion is required and dogs, like humans, will
generate only enough to literally fall forward, catching their weight
with each stride. At a trot however, greater speed is required, as well
as lift, and the propelling leg must generate more power. At the support
position, the hindquarter will sink slightly, increasing angulation
and the legs potential thrust . A dog with very little angulation
can produce very little thrust at the trot (Fig 9). Ideal angulation
produces the most thrust for the least expenditure of energy. The over
angulated dog can often produce good thrust, but he must also expend
more energy to forcefully extend his overangulated legs when in motion,
and to keep the acutely angled levers of his hind legs from collapsing
when in the support position. Because about half the total length of
the stride is actually involved in generating power, total length of
stride is important. The greater the angulation the longer the stride,
but the cost in energy consumption goes up.



As the hind leg is thrusting backwards,
several things are happening (Fig 10). The femur or thigh bone is moving
backwards, the result of powerful contractions of the muscles connecting
the rear pelvis and thigh. The knee is straightening, because of the
contractions of the muscles attached to the femur and shin, over the
patella. The patella adds a bit of extra distance the tendon must stretch,
increasing leverage. Finally, the hock joint is straightening, because
of the action of the calf muscle previously discussed. All these massive
muscles work in concert to produce a single, powerful, straightening
thrust of the leg. In a dog lacking angulation, these muscles are shorter
and cannot contract to the same degree as those of a well angulated
dog, producing less thrust. More than muscle work is involved however.
At the support position the leg is well bent, and the tendons running
over the knee and the hock joint (the Achilles) are stretched like elastics,
storing energy (Fig 11). As the leg moves backwards, this energy is
released as the tendons snap back to their normal lengths, further helping
to open the knee and hock joints. The overangulated dog, with over bent
joints, has overlong, sloppy tendons that have limited ability to stretch
and store energy. Also, as the foot moves backwards, the toes are stretched
back, much like bending your fingers back, stretching their tendons
and adding even more snap to the stride (Fig 12). Longer toes that are
well knuckled up give the most leverage and elastic snap. Flat feet
give the least. The propulsive power of short, strong tendons is free
power, in the sense that it requires no muscular work, and is the best
argument against overangulation.
At
the end of the stride the leg will be nearly, but not completely straight
(Fig 13). Not completely, because the trot is not a gait of maximum
exertion. The dog that shows a maximum degree of stride length at the
trot is expending too much energy, and while it will no doubt show a
visually spectacular side gait, it cannot keep this up for a prolonged
period of time. At increased speeds, the gallop is more cost efficient.
Also, extreme side gaiters tend to show diminished follow-through, with
the hind foot leaving the ground but failing to continue its sweep backwards.
Follow-through is important, much like the back swing of a golfers
club. It finishes the stride, allowing the thrust to die gradually and
ensuring that the most powerful part of the swing or stride is expended
where it will do the most good; on the ground. The extreme mover will
often travel so low to the ground that its toes are dragging in the
grass. It is probably better that a dog kick up somewhat behind, than
show no follow-through at all.

At
the same time that one leg is thrusting backwards, the other rear leg
is moving ahead (Fig 14). It should move low to the ground. Hoisting
a heavy leg higher than necessary uses energy. Having to reach too far
ahead also takes energy. Most dogs, whatever their structure, attempt
to reach to their centre of gravity, which is usually located a bit
behind the centre of the rib cage, to find their balance. A correctly
structured dog need not reach so far ahead as a long bodied dog. When
the foot has found the ground, it begins to support the weight of the
body being propelled over it, and to fold into the support position
in preparation for the next stride. It should be noted that the rear
foot should strike the ground with the plantar pad first, and the toes
second, but never with the heel or hock. Overangulated dogs, especially
those with sickle hocks, tend to show a flat footed stride, forcing
the hock to absorb a ground strike for which it is not designed (Fig
15). If you examine the bottom of the dogs hock and see what appears
to be an extra pad of callused skin half way up, or worse, at the tip
of the hock, this is due to the hocks repeated abrasion with the
ground and is unacceptable.